Lancer bears the look of a typical medieval knight, wearing full plate armour, etched wih leaf designs on the helmet, pauldrons and cuirass. He carries a relatively short, broad lance in his right hand and his left vambrace seems to have an ornate, many pointed shield integrated with it.
Lancer seemed to enjoy combat, becoming excited at the prospect of fighting Saber, even complimenting her in her near instantaneous victory over him. Despite this, he seems to have little respect for non-combatants, not hesitating to fulfill orders to kill two teenagers before him, and referring to one as a "damn monkey" when fleeing from him.
On a train, his Master the Mysterious Officer orders him to kill both Tsukumo Fujimiya and the newly arrived Kanata Akagi. He chases them through the train and catches up just as Tsukumo attempts to summon a Servant. Kanata tackles her away from his spear strike, but the spear breaks open Kanata's backpack and a light novel falls into the magic circle, resulting in Saber being summoned. He is impressed when Saber dodges his spear and attempts to introduce himself, only for Saber to stab him in the throat mid-sentence. As he staggers around choking on his own blood and marveling at how fast and precise her swordsmanship is to be able to exploit the gaps in his armor, Saber finishes him off and calls him a fool for trying to talk in the middle of a fight. Later, Saber hurls his helmet at Archer to try to distract her.
Despite not being a true servant, Lancer still possesses abilities far beyond an average human. With a thrust of his lance he created multiple beams of light which eradicated the wall of a train, extending several feet into the carriage to destroy Kanata Akagi's backpack.
Lancer takes the place of Cú Chulainn in the adaptation.